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On August 24, 2007, the portion of the highway between Glen Miller Road in Trenton and the Don Valley Parkway / Highway 404 Junction in Toronto was designated the Highway of Heroes, as the road is travelled by funeral convoys for fallen Canadian Forces personnel from CFB Trenton to the coroner's office in Toronto.On September 27, 2013, the Highway of Heroes designation was extended west to Keele Street in Toronto, to coincide with the move of the coroner's office to the new Forensic Services and Coroner's Complex at the Humber River Hospital.To the south of Woodstock, Highway 401 curves northeast and encounters the western terminus of Highway 403.After this 10 km (6.2 mi) gap, the highway interchanges with the Highway 407 Express Toll Route.

East of the interchange with Renforth Drive, the collector lanes diverge to become the on-ramps to Highway 427 northbound and southbound.In 1965 it was given a second designation, the Macdonald–Cartier Freeway, in honour of two Fathers of Confederation.At the end of 1968, the Gananoque–Brockville section was bypassed and the final intersection grade-separated near Kingston, making Highway 401 a freeway for its entire 817.9-km length.In addition, Highway 401 was equipped with a traffic camera system called COMPASS in early 1991.Using closed-circuit television cameras, vehicle detection loops and LED changeable-message signs, COMPASS enables the MTO Traffic Operations Centre to obtain a real-time assessment of traffic conditions and alert drivers of collisions, congestion and construction.

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